Some holoenzymes, such as pyruvate dehydrogenase, are highly compex, with several cofactors.

有一些全酶是高度复杂，如丙酮酸脱氢酶。

In working skeletal muscle under anaerobic conditions, glyceraldehyde 3-phosphate is converted to pyruvate (the payoff phase of glycolysis), and the pyruvate is reduced to lactate .

骨骼肌在厌件下工作，3-磷酸甘油转变成丙酮酸(糖收获阶段)，丙酮酸又转变成乳酸。