Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足迹同样是常用计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量老虎脚印长度和宽度以及脚趾之间角度，以及10种其他变量来确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足迹同样是常用计算老虎数方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片老虎脚印长和宽脚趾之间角，10种其他变确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足迹同样是常用计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量老虎脚印长度和宽度以及脚趾之间角度，以及10种其他变量来确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足迹同样是常用计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量老虎脚印长度和宽度以及脚趾之间角度，以及10种其他变量来确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足迹同样是常用计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量老虎脚印长度和宽度以及脚趾之间角度，以及10种其他变量来确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足迹同样是常用计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量老虎脚印长度和宽度以及脚趾之间角度，以及10种其他变量来确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足迹同样是常计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人些照片来测量老虎脚印长度和宽度以及脚趾之度，以及10种其他变量来确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎同样是常用计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量老虎脚印长度和宽度及脚趾之间角度，及10种其他变量来确定不同老虎“身份”。

Paw print, or pugmark, tracing is another common technique for counting tigers.

爪印和老虎足是常用计算老虎数量方法。

The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.

研究人员用这些照片来测量老虎脚印长度和宽度以及脚趾之间角度，以及10种其他变量来确定不老虎“”。